Bombus Latreille, 1802
(bumble bees or humble bees & cuckoo bees)

Interactions where Bombus is the dominant partner and gains from the process

The following relationships have been collated from the published literature (see 'Interaction References').

Stage Summary Taxon Vernacular Classification References Darwin Classification of Active Taxon Active Taxon Active Vernacular Active Taxon Uncertain Active State Active Part Active Stage Relationship Relationship Uncertain Relationship Geography Darwin Classification of Passive Taxon Passive Taxon Passive Vernacular Passive Taxon Uncertain Passive State Passive Part Passive Stage Passive Taxon's significance to Active Taxon Indoors etc Season
queen queen pollinates or fertilises flower Major Anacamptis morio Green-winged Orchid Asparagales: Orchidaceae Orchids of Britain & Ireland: a field and site guide, Harrap, A. & S., 2009
/Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesqueen Plant / pollinated /Plantae/Tracheophyta/Magnoliopsida/Asparagales/Orchidaceae/Anacamptis morio/Anacamptis moriogreen-winged orchidflowerMajor
adult adult pollinates or fertilises flower Dactylorhiza incarnata Early Marsh-orchid Asparagales: Orchidaceae Orchids of Britain & Ireland: a field and site guide, Harrap, A. & S., 2009
/Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesadult Plant / pollinated /Plantae/Tracheophyta/Magnoliopsida/Asparagales/Orchidaceae/Dactylorhiza incarnata/Dactylorhiza incarnataearly marsh-orchidflower
adult adult pollinates or fertilises live flower Epipogium aphyllumQuestionable Ghost Orchid Asparagales: Orchidaceae Orchids of Britain & Ireland: a field and site guide, Harrap, A. & S., 2009
/Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesadult Plant / pollinated /Plantae/Tracheophyta/Magnoliopsida/Asparagales/Orchidaceae/Epipogium aphyllum/Epipogium aphyllumghost orchid?liveflower
adult adult pollinates or fertilises flower Goodyera repens Creeping Lady’s-tresses Asparagales: Orchidaceae Orchids of Britain & Ireland: a field and site guide, Harrap, A. & S., 2009
/Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesadult Plant / pollinated /Plantae/Tracheophyta/Magnoliopsida/Asparagales/Orchidaceae/Goodyera repens/Goodyera repenscreeping lady's-tressesflower
adult adult pollinates or fertilises flower Major Orchis mascula Early Purple Orchid, Early-purple Orchid Asparagales: Orchidaceae Orchids of Britain & Ireland: a field and site guide, Harrap, A. & S., 2009
/Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesadult Plant / pollinated /Plantae/Tracheophyta/Magnoliopsida/Asparagales/Orchidaceae/Orchis mascula/Orchis masculaearly purple orchid, early-purple orchidflowerMajor
adult adult pollinates or fertilises flower Orchis militaris Soldier Orchid, Military Orchid Asparagales: Orchidaceae Orchids of Britain & Ireland: a field and site guide, Harrap, A. & S., 2009
/Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesadult Plant / pollinated /Plantae/Tracheophyta/Magnoliopsida/Asparagales/Orchidaceae/Orchis militaris/Orchis militarissoldier orchid, military orchidflower
adult adult pollinates or fertilises flower Spiranthes spiralis Autumn Lady’s-tresses Asparagales: Orchidaceae Orchids of Britain & Ireland: a field and site guide, Harrap, A. & S., 2009
/Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesadult Plant / pollinated /Plantae/Tracheophyta/Magnoliopsida/Asparagales/Orchidaceae/Spiranthes spiralis/Spiranthes spiralisautumn lady's-tressesflower

Interactions where Bombus is the victim or passive partner (and generally loses out from the process)

The following relationships have been collated from the published literature (see 'Interaction References').

Filters:

Affected Part Summary Taxon Vernacular Classification References Darwin Classification of Active Taxon Active Taxon Active Vernacular Active Taxon Uncertain Active State Active Part Active Stage Relationship Relationship Uncertain Relationship Geography Darwin Classification of Passive Taxon Passive Taxon Passive Vernacular Passive Taxon Uncertain Passive State Passive Part Passive Stage Passive Taxon's significance to Active Taxon Indoors etc Season Summary
adult is predated by Foreign Merops apiaster Bee-eater, European Bee-eater Coraciiformes: Meropidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Chordata/Aves/Coraciiformes/Meropidae/Merops apiaster/Merops apiasterbee-eater, european bee-eater Animal / predatorForeign /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesadult is predator of adult
adult is predated by Lanius shrikes Passeriformes: Laniidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Chordata/Aves/Passeriformes/Laniidae/Lanius/Laniusshrikes Animal / predator /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesadult is predator of adult
adult is predated by Major Lanius collurio Red-backed Shrike Passeriformes: Laniidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Chordata/Aves/Passeriformes/Laniidae/Lanius collurio/Lanius colluriored-backed shrike Animal / predator /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesadultMajor is predator of adult
adult is predated by UK/Ireland Muscicapa striata Spotted Flycatcher Passeriformes: Muscicapidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Chordata/Aves/Passeriformes/Muscicapidae/Muscicapa striata/Muscicapa striataspotted flycatcher Animal / predatorUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesadult is predator of adult
nest is predated by Meles meles Badger, Eurasian Badger Carnivora: Mustelidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Chordata/Mammalia/Carnivora/Mustelidae/Meles meles/Meles melesbadger, eurasian badger Animal / predator /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesnest is predator of nest
nest is predated by Foreign Neovison vison American Mink Carnivora: Mustelidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Chordata/Mammalia/Carnivora/Mustelidae/Neovison vison/Neovison visonAmerican mink Animal / predatorForeign /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesnest is predator of nest
queen may give a ride to female UK/IrelandQuestionable Scutacarus a genus of mites Trombidiformes: Scutacaridae Bee Mite ID: Bee-Associated Mite Genera of the World, Klimov, P.B., O'Connor, B., Ochoa, R., Bauchan, G., Redford, A. & Scher, J., 2016
Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Arthropoda/Arachnida/Trombidiformes/Scutacaridae/Scutacarus/Scutacarusa genus of mitesfemale Animal / phoresyRelationship uncertainUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesqueen female hitches a ride on queen
adult may give a ride to adult UK/Ireland Parasitellus fucorum Bumble Bee Mites Mesostigmata: Parasitidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Arthropoda/Arachnida/Mesostigmata/Parasitidae/Parasitellus fucorum/Parasitellus fucorumbumble bee mitesadult Animal / phoresyUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesadult adult hitches a ride on adult
is predated by UK/Ireland ARANEAE spiders Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Arthropoda/Arachnida/Araneae/Araneaespiders Animal / predatorUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo bees is predator of
nest animal debris may support larva UK/Ireland Volucella bombylans Fool’s Bumble-bee Diptera: Syrphidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Arthropoda/Insecta/Diptera/Syrphidae/Volucella bombylans/Volucella bombylansfool's bumble-beelarva Animal / dung / debris saprobeUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesnest larva feeds on animal debris nest
adult is endoparasitoid host of larva UK/Ireland CONOPIDAE thick-headed flies, or beegrabbers and waspgrabbers Diptera Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Arthropoda/Insecta/Diptera/Conopidae/Conopidaethick-headed flies, or beegrabbers and waspgrabberslarva Animal / parasitoid / endoparasitoidUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesadult larva is endoparasitoid of adult
nest may attract larva Fannia canicularis Latrine Fly, Small House-fly, Lesser House-fly Diptera: Fanniidae Diptera: Cyclorrhapha Calyptrata (I) Sect (a) Tachinidae and Calliphoridae, van Emden, F.I.,, 1954
/Animalia/Arthropoda/Insecta/Diptera/Fanniidae/Fannia canicularis/Fannia canicularislatrine fly, small house-fly, lesser house-flylarva Animal / associate /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesnest larva is associated with nest
nest may attract larva Fannia scalaris a lesser house fly Diptera: Fanniidae Diptera: Cyclorrhapha Calyptrata (I) Sect (a) Tachinidae and Calliphoridae, van Emden, F.I.,, 1954
/Animalia/Arthropoda/Insecta/Diptera/Fanniidae/Fannia scalaris/Fannia scalarisa lesser house flylarva Animal / associate /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesnest larva is associated with nest
prepupa is ectoparasitoid host of larva UK/Ireland Brachicoma devia a flesh fly Diptera: Sarcophagidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
Diptera: Cyclorrhapha Calyptrata (I) Sect (a) Tachinidae and Calliphoridae, van Emden, F.I.,, 1954
/Animalia/Arthropoda/Insecta/Diptera/Sarcophagidae/Brachicoma devia/Brachicoma deviaa flesh flylarva Animal / parasitoid / ectoparasitoidUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesprepupa larva is ectoparasitoid of prepupa
nest accommodates puparium UK/Ireland Brachicoma devia a flesh fly Diptera: Sarcophagidae Diptera: Cyclorrhapha Calyptrata (I) Sect (a) Tachinidae and Calliphoridae, van Emden, F.I.,, 1954
/Animalia/Arthropoda/Insecta/Diptera/Sarcophagidae/Brachicoma devia/Brachicoma deviaa flesh flypuparium Animal / resting place / withinUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesnest puparium rests inside nest
soil near nest accommodates puparium UK/Ireland Brachicoma devia a flesh fly Diptera: Sarcophagidae Diptera: Cyclorrhapha Calyptrata (I) Sect (a) Tachinidae and Calliphoridae, van Emden, F.I.,, 1954
/Animalia/Arthropoda/Insecta/Diptera/Sarcophagidae/Brachicoma devia/Brachicoma deviaa flesh flypuparium Animal / resting place / withinUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beessoil near nest puparium rests inside soil near nest
nest may attract larva Macronychia griseola a flesh fly Diptera: Sarcophagidae Diptera: Cyclorrhapha Calyptrata (I) Sect (a) Tachinidae and Calliphoridae, van Emden, F.I.,, 1954
/Animalia/Arthropoda/Insecta/Diptera/Sarcophagidae/Macronychia griseola/Macronychia griseolaa flesh flylarva Animal / associate /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesnest larva is associated with nest
nest may attract larva Macronychia polyodon a flesh fly Diptera: Sarcophagidae Diptera: Cyclorrhapha Calyptrata (I) Sect (a) Tachinidae and Calliphoridae, van Emden, F.I.,, 1954
/Animalia/Arthropoda/Insecta/Diptera/Sarcophagidae/Macronychia polyodon/Macronychia polyodona flesh flylarva Animal / associate /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesnest larva is associated with nest
nest is parasitised by female UK/Ireland Mutilla europaea Large Velvet Ant Hymenoptera: Mutillidae Forage, prey and host-parasite relationships of aculeates, Macdonald, M.A., 2007
/Animalia/Arthropoda/Insecta/Hymenoptera/Mutillidae/Mutilla europaea/Mutilla europaealarge velvet antfemale Animal / parasiteUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesnest female parasitises nest
nest animal debris may support silken tube inhabiting, young, colonial larva UK/Ireland Aphomia sociella Bee Moth Lepidoptera: Pyralidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Arthropoda/Insecta/Lepidoptera/Pyralidae/Aphomia sociella/Aphomia sociellabee mothsilken tube inhabiting, young, coloniallarva Animal / dung / debris saprobeUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesnest silken tube inhabiting, young, colonial larva feeds on animal debris nest
food stores is kleptoparasitised by silken tube inhabiting, older, colonial larva UK/Ireland Aphomia sociella Bee Moth Lepidoptera: Pyralidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Arthropoda/Insecta/Lepidoptera/Pyralidae/Aphomia sociella/Aphomia sociellabee mothsilken tube inhabiting, older, coloniallarva Animal / kleptoparasiteUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesfood stores silken tube inhabiting, older, colonial larva kleptoparasitises food stores
cells is predated by silken tube inhabiting, older, colonial larva UK/Ireland Aphomia sociella Bee Moth Lepidoptera: Pyralidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Arthropoda/Insecta/Lepidoptera/Pyralidae/Aphomia sociella/Aphomia sociellabee mothsilken tube inhabiting, older, coloniallarva Animal / predatorUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beescells silken tube inhabiting, older, colonial larva is predator of cells
larvae is predated by silken tube inhabiting, older, colonial larva UK/Ireland Aphomia sociella Bee Moth Lepidoptera: Pyralidae Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Arthropoda/Insecta/Lepidoptera/Pyralidae/Aphomia sociella/Aphomia sociellabee mothsilken tube inhabiting, older, coloniallarva Animal / predatorUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beeslarvae silken tube inhabiting, older, colonial larva is predator of larvae
abdomen abdomen of queen is endoparasitised by adult female UK/Ireland Sphaerularia bombi Bumblebee Roundworm Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Nematoda/Sphaerularia bombi/Sphaerularia bombibumblebee roundwormadult female Animal / parasite / endoparasiteUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesabdomenqueen (Spring)adult female endoparasitises abdomen of queen
blood blood of queen is endoparasitised by larva UK/Ireland Sphaerularia bombi Bumblebee Roundworm Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Nematoda/Sphaerularia bombi/Sphaerularia bombibumblebee roundwormlarva Animal / parasite / endoparasiteUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesbloodqueen (Spring)larva endoparasitises blood of queen
frass frass of queen may give a ride to larva UK/Ireland Sphaerularia bombi Bumblebee Roundworm Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Nematoda/Sphaerularia bombi/Sphaerularia bombibumblebee roundwormlarva Animal / phoresyUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesfrassqueen (Spring)larva hitches a ride on frass of queen
gut gut of queen is endoparasitised by larva UK/Ireland Sphaerularia bombi Bumblebee Roundworm Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Nematoda/Sphaerularia bombi/Sphaerularia bombibumblebee roundwormlarva Animal / parasite / endoparasiteUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesgutqueen (Spring)larva endoparasitises gut of queen
reproductive system reproductive system of queen is endoparasitised by larva UK/Ireland Sphaerularia bombi Bumblebee Roundworm Bumblebees, Prys-Jones, O.E. & Corbet, S.A., 1987
/Animalia/Nematoda/Sphaerularia bombi/Sphaerularia bombibumblebee roundwormlarva Animal / parasite / endoparasiteUK and/or Eire /Animalia/Arthropoda/Insecta/Hymenoptera/Apidae/Bombus/Bombusbumble bees or humble bees & cuckoo beesreproductive systemqueen (Spring)larva endoparasitises reproductive system of queen

Author & YearTitleSource
Harrap, A. & S., 2009Orchids of Britain & Ireland: a field and site guide2nd edition, 480pp, A&C Black Publishers Ltd
Klimov, P.B., O'Connor, B., Ochoa, R., Bauchan, G., Redford, A. & Scher, J., 2016Bee Mite ID: Bee-Associated Mite Genera of the Worldidtools.org/id/mites/beemites
Macdonald, M.A., 2007Forage, prey and host-parasite relationships of aculeates
Prys-Jones, O.E. & Corbet, S.A., 1987BumblebeesNaturalists’ Handbooks, 6, 92pp, The Richmond Publishing Co. Ltd
van Emden, F.I.,, 1954Diptera: Cyclorrhapha Calyptrata (I) Sect (a) Tachinidae and CalliphoridaeHandbooks for the Identification of British Insects , Vol X. Part 4(a), 133pp, The Royal Entomological Society of London

Bombus (bumble bees or humble bees & cuckoo bees) may also be included in 'feeds on' relations listed under the following higher taxa:

Bombus (bumble bees or humble bees & cuckoo bees) may also be included in 'fed on by' relations listed under the following higher taxa:

Subtaxon Rank Featured
subtaxa
No of
interactions
No of
references
Subgenus 13 subtaxa 30 trophisms 14 references
Subgenus 6 subtaxa 11 trophisms 4 references
Species 1 trophisms
Species 1 trophisms
Taxonomic hierarchy:
GenusBombus (bumble bees or humble bees & cuckoo bees)
FamilyAPIDAE (bees)
SuperfamilyAPOIDEA (solitary and hive bees)
InfraorderAculeata (aculeates: bees, wasps and ants)
SuborderAPOCRITA (bees, wasps, ants and parasitoids)
OrderHYMENOPTERA (ants, bees and wasps, sawflies and parasitoid wasps)
Division Endopterygota (bees, beetles, flies, moths and other insects with wings developing internally)
InfraclassNeoptera (bees, beetles, flies, grasshoppers, moths and other advanced insects)
SubclassPTERYGOTA (bees, beetles, dragonflies, flies, grasshoppers, moths and other winged insects)
ClassINSECTA (true insects)
SubphylumHEXAPODA (insects and other 6-legged organisms)
PhylumARTHROPODA (arthropods)
SuperphylumECDYSOZOA (skin shedders)
CladeBilateria (bilaterally symmetrical animals)
SubkingdomEUMETAZOA (metazoans)
KingdomANIMALIA (animals)
DomainEukaryota (eukaryotes)
LifeBIOTA (living things)
NBNNBN (data.nbn.org.uk) has a distribution map for Bombus (bumble bees or humble bees & cuckoo bees)

Identification Works

Handling & MagnificationAuthorYearTitleSource
Anon. Bumblebee pages www.bumblebee.org
Benton, E. 2006 Bumblebees New Naturalists, 98, Collins
Edwards, M. & Jenner, M. 2005 Field Guide to Bumblebees of Great Britain and Ireland 108pp
in the field () Falk, S. 2011 Warwickshire’s bumblebees
in the field () Prys-Jones, O.E. & Corbet, S.A. 1987 Bumblebees Naturalists’ Handbooks, 6, 92pp, The Richmond Publishing Co. Ltd
Ptácek, V. & Zaludová, R. Bumble bees www.sci.muni.cz/ptacek/bombi.htm

Bombus (bumble bees or humble bees & cuckoo bees) may also be included in identification literature listed under the following higher taxa:

General Works

AuthorYearTitleSource
Anon. Bumblebee pages www.bumblebee.org
Edwards, M. & Williams, P. 2004 Where have all the bumblebees gone, and could they ever return? British Wildlife Vol 15, No. 5: 305-312.
Heath, J. & Alford, D.V. 1973 Provisional atlas of the insects of the British Isles. part 3 Hymenoptera - Apidae (Bombus:Psithyrus) Bumblebees. 27pp, Biological Records Centre

Parasites

Colla, S.R., Otterstatter, M.C., Gegear, R.J. & Thomson, J.D. 2006 Plight of the bumble bee: Pathogen spillover from commercial to wild populations Biological Conservation Vol 129: 461-467.
Korner, P. & Schmid-Hempel, P. 2005 Correlates of parasite load in bumblebees in an Alpine habitat Entomological Science Vol 8 (2): 151-160.

Pathology

Klimov, P.B., O’Connor, B., Ochoa, R., Bauchan, G., Redford, A. & Scher, J. 2016 Bee Mite ID: Bee-Associated Mite Genera of the World idtools.org/id/mites/beemites

Literature listed under the following higher taxa may also be relevant to Bombus (bumble bees or humble bees & cuckoo bees):

BioImagesBioImages (www.bioimages.org.uk) has 232 images of Bombus (bumble bees or humble bees & cuckoo bees)
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